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55. Boulay R, X Arnan, X Cerdá, J Retana 2014. The ecological benefits of larger colony size may promote polygyny in ants. Journal of evolutionary biology 27: 2856-2863


54. Cerdá X., L Van Oudenhove, C Bernstein, R Boulay 2014. A list of and some comments about the trail pheromones of ants. Natural product communications 9: 1115-1122


53. Lenoir A, Touchard A, Devers S, Christidès JP, Boulay R, Cuvillier-Hot V 2014 Ant cuticular response to phthalate pollution. Environmental Science and Pollution Research 21: 13446-13451


52. Jowers MJ, Amor F, Ortega P, Boulay R, Cerdá X, Lenoir A, Galarza JA. 2014 Recent allopatric speciation and secondary contact in endemic ants. Molecular Ecology 23: 2529-2542


51. Cuvillier-Hot V, Salin K, Devers S, Tasiemski A, Schaffner P, Boulay R, Billard S, Lenoir A 2014 Impact of ecological doses of the most widespread phthalate on a terrestrial species, the ant Lasius niger. Environmental Research 131, 104-110.


50. Caut S, MJ Jowers, X Arnan, J Pearce‐Duvet, A Rodrigo, X Cerdá, R Boulay. 2014. The effects of fire on ant trophic assemblage and sex allocation. Ecology and Evolution 4, 35-49.


49. Lenoir A, J Háva, A Hefetz, A Dahbi, X Cerdá, R Boulay. 2013. Chemical integration of Thorictus myrmecophilous beetles into Cataglyphis ant nests. Biochemical Systematics and Ecology 51, 335-342


48. Jowers MJ, L Leniaud, X Cerdá, S Alasaad, S Caut, F Amor, S Aron, R Boulay. 2013. Social and population structure in the ant Cataglyphis emmae. PloS one 8, e72941


47. Lázaro‐González A, X Arnan, R Boulay, X Cerdá, A Rodrigo. 2013. Short‐term ecological and behavioural responses of Mediterranean ant species Aphaenogaster gibbosa (Latr. 1798) to wildfire. Insect Conservation and Diversity 6, 627-638


46. Caut S, MJ Jowers, X Cerdá, R Boulay. 2013. Questioning the mutual benefits of myrmecochory: a stable isotope‐based experimental approach. Ecological Entomology 38: 390-399


45. Caut S, Á Barroso, X Cerdá, F Amor, R Boulay. 2013. A year in an ant's life: opportunism and seasonal variation in the foraging ecology of Aphaenogaster senilis. Ecoscience 20, 19-27


44. Ruel C, A Hefetz, X Cerdá, R Boulay. 2013. Recognition of caste and mating status maintains monogyny in the ant Aphaenogaster senilis Behavioral Ecology and Sociobiology 67, 1295-1305


43. Ruel C, A Lenoir, X Cerdá, R Boulay. 2013. Surface lipids of queen-laid eggs do not regulate queen production in a fission-performing ant. Naturwissenschaften 100, 91-100


42. Barroso Á, F Amor, X Cerdá, R Boulay. 2013. Dispersal of non-myrmecochorous plants by a “keystone disperser” ant in a Mediterranean habitat reveals asymmetric interdependence. Insectes Sociaux 60, 75-86


41. Ruel C, X Cerdá, R Boulay. 2012. Behaviour-mediated group size effect constrains reproductive decisions in a social insect. Animal Behaviour 84, 853-860


40. Leniaud L, H Darras, R Boulay, S Aron. 2012. Social hybridogenesis in the clonal ant Cataglyphis hispanica. Current Biology 22, 1188-1193


39. van Oudenhove L, R Boulay, A Lenoir, C Bernstein, X Cerdá 2012. Substrate Temperature Constrains Recruitment and Trail Following Behavior in Ants. Journal of Chemical Ecology 38, 802-809


38. Lenoir A, Q Chalon, A Carvajal, C Ruel, Á Barroso, T Lackner, R Boulay. 2012. Chemical integration of myrmecophilous guests in Aphaenogaster ant nests. Psyche: A Journal of Entomology


37. Galarza JA, R Jovani, X Cerdá, C Rico, Á Barroso, R Boulay. 2012. Frequent colony relocations do not result in effective dispersal in the gypsy ant Aphaenogaster senilis. Oikos 121, 605-613


36. van Oudenhove L, E Billoir, R Boulay, C Bernstein, X Cerdá 2011. Temperature limits trail following behaviour through pheromone decay in ants. Naturwissenschaften 98, 1009-1017


35. Amor F, P Ortega, MJ Jowers, X Cerdá, J Billen, A Lenoir, R Boulay. 2011. The evolution of worker–queen polymorphism in Cataglyphis ants: interplay between individual-and colony-level selections. Behavioral Ecology and Sociobiology 65, 1473-1482


34. Lenoir A, A Benoist, A Hefetz, W Francke, X Cerdá, R Boulay. 2011. Trail-following behaviour in two Aphaenogaster ants. Chemoecology 21, 83-88.


33. Amor F, P Ortega, X Cerdá, R Boulay. 2011. Solar elevation triggers foraging activity in a thermophilic ant. Ethology 117, 1031-1039.


32. Boulay R, JA Galarza, B Chéron, A Hefetz, A Lenoir, L Oudenhove, X Cerdá. 2010. Intraspecific competition affects population size and resource allocation in an ant dispersing by colony fission. Ecology 91, 3312-3321.


31. Dejean A, C Leroy, B Corbara, R Céréghino, O Roux, B Hérault, V Rossi, R Boulay. 2010. A temporary social parasite of tropical plant-ants improves the fitness of a myrmecophyte. Naturwissenschaften 97, 925-934.


30. Dejean A, C Leroy, B Corbara, O Roux, R Céréghino, J Orivel, R Boulay. 2010. Arboreal ants use the “Velcro® principle” to capture very large prey. PloS one 5 (6), e11331.


29. Amor F, P Ortega, X Cerdá, R Boulay 2010. Cooperative prey-retrieving in the ant Cataglyphis floricola: an unusual short-distance recruitment. Insectes Sociaux 57, 91-94.


28. El Keroumi A, K Naamani, A Dahbi, I Luque, A Carvajal, X Cerda, R Boulay. 2010. Effect of ant predation and abiotic factors on the mortality of medfly larvae, Ceratitis capitata, in the Argan forest of Western Morocco. Biocontrol Science and Technology 20, 751-762.


27. Ichinose K, R Boulay, X Cerdá, A Lenoir 2009. Influence of queen and diet on nestmate recognition and cuticular hydrocarbon differentiation in a fission-dispersing ant, Aphaenogaster senilis. Zoological Science 26, 681-685.


26. Boulay R, X Cerdá, A Fertin, K Ichinose, A Lenoir 2009. Brood development into sexual females depends on the presence of a queen but not on temperature in an ant dispersing by colony fission, Aphaenogaster senilis. Ecological Entomology 34, 595-602.


25. Boulay R, F Carro, RC Soriguer, X Cerdá 2009. Small-scale indirect effects determine the outcome of a tripartite plant–disperser–granivore interaction. Oecologia 161, 529-537.


24. Galarza JA, R Boulay, X Cerdá, C Doums, P Federici, H Magalon, T Monnin, C. Rico. 2009. Development of single sequence repeat markers for the ant Aphaenogaster senilis and cross-species amplification in A. iberica, A. gibbosa, A. subterranea and Messor maroccanus. Conservation Genetics 10, 519-521.


23. Cerdá X, E Angulo, R Boulay, A Lenoir. 2009. Individual and collective foraging decisions: a field study of worker recruitment in the gypsy ant Aphaenogaster senilis. Behavioral Ecology and Sociobiology 63, 551-562.


22. Katzav-Gozansky T, R Boulay, A Ionescu-Hirsh, A Hefetz 2008. Nest volatiles as modulators of nestmate recognition in the ant Camponotus fellah. Journal of insect physiology 54, 378-385.


21. Boulay R, X Cerdá, T Simon, M Roldan, A Hefetz 2007. Intraspecific competition in the ant Camponotus cruentatus: should we expect the dear enemy effect? Animal Behaviour 74, 985-993


20. Boulay R, F Carro, RC Soriguer, X Cerdá 2007. Synchrony between fruit maturation and effective dispersers' foraging activity increases seed protection against seed predators. Proceedings of the Royal Society B: Biological Sciences 274, 2515-2522.


19. Manzaneda A, PJ Rey, R Boulay. 2007. Geographic and temporal variation in the ant–seed dispersal assemblage of the perennial herb Helleborus foetidus L. (Ranunculaceae). Biological Journal of the Linnean Society 92, 135-150.


18. Boulay R, A Hefetz, X Cerdá, S Devers, W Francke, R Twele, A Lenoir 2007. Production of sexuals in a fission-performing ant: dual effects of queen pheromones and colony size. Behavioral Ecology and Sociobiology 61, 1531-1541.


17. Alcántara JM, PJ Rey, AJ Manzaneda, R Boulay, JM Ramírez, JM Fedriani. 2007. Geographic variation in the adaptive landscape for seed size at dispersal in the myrmecochorous Helleborus foetidus. Evolutionary Ecology 21, 411-430.


16. Boulay R, J Coll-Toledano, AJ Manzaneda, X Cerdá 2007. Geographic variations in seed dispersal by ants: are plant and seed traits decisive? Naturwissenschaften 94, 242-246.


15. Fedriani JM, R Boulay 2006. Foraging by fearful frugivores: combined effect of fruit ripening and predation risk. Functional Ecology 20, 1070-1079.


14. Boulay R, J Coll-Toledano, X Cerdá. 2006. Geographic variations in Helleborus foetidus elaiosome lipid composition: implications for dispersal by ants. Chemoecology 16, 1-7.


13. Turillazzi S, L Dapporto, C Pansolli, R Boulay, FR Dani, G Moneti 2006. Habitually used hibernation sites of paper wasps are marked with venom and cuticular peptides. Current biology, R530-R5311.


12. Boulay R, JM Fedriani, AJ Manzaneda, X Cerdá 2005.Indirect effects of alternative food resources in an ant–plant interaction. Oecologia 144, 72-79.


11. Katzav-Gozansky T, R Boulay, V Soroker, A Hefetz 2004. Queen–signal modulation of worker pheromonal composition in honeybees. Proceedings of the Royal Society of London. Series B: Biological Sciences 271, 2065-2069.


10. Katzav-Gozansky T, R Boulay, R Vander Meer, A Hefetz 2004. In-nest environment modulates nestmate recognition in the ant Camponotus fellah. Naturwissenschaften 91, 186-190.


9. Boulay R, T Katzav-Gozansky, A Hefetz, A Lenoir 2004. Odour convergence and tolerance between nestmates through trophallaxis and grooming in the ant Camponotus fellah (Dalla Torre). Insectes sociaux 51, 55-61.


8. Boulay R, T Katzav-Gozansky, RK Vander Meer, A Hefetz 2003 Colony insularity through queen control on worker social motivation in ants. Proceedings of the Royal Society of London. Series B: Biological Sciences 270, 971-977.


7. Boulay R, LM Hooper-Bui 2002. The effect of chlordimeform on the foraging activity of the red imported fire ant Solenopsis invicta (Hymenoptera: Formicidae). Sociobiology 39, 129-139


6. Boulay R, LM Hooper‐Bui, J Woodring 2001. Oviposition and oogenesis in virgin fire ant females Solenopsis invicta are associated with a high level of dopamine in the brain. Physiological Entomology 26, 294-299.


5. Boulay R, A Lenoir. 2001 Social isolation of mature workers affects nestmate recognition in the ant Camponotus fellah. Behavioural processes 55, 67-73.


4. Boulay R, V Soroker, EJ Godzinska, A Hefetz, A Lenoir 2000. Octopamine reverses the isolation-induced increase in trophallaxis in the carpenter ant Camponotus fellah. Journal of Experimental Biology 203, 513-520.


3. Auger J, R Boulay, B Jaillais, S Delion-Vancassel 2000. Analysis of biogenic amines by solid-phase microextraction and high-performance liquid chromatography with electrochemical detection. Journal of Chromatography A 870, 395-403.


2. Boulay R, A Hefetz, V Soroker, A Lenoir 2000. Camponotus fellah colony integration: worker individuality necessitates frequent hydrocarbon exchanges. Animal Behaviour 59, 1127-1133.


1. Boulay R, M Quagebeur, EJ Godzinska, A Lenoir 1999. Social isolation in ants: evidence of its impact on survivorship and behavior in Camponotus fellah (Hymenoptera: Formicidae). Sociobiology 33, 111-124.

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